Analysing the effect of stand density and site conditions on structure and growth of oak species using Nelder trials along an environmental gradient: experimental design, evaluation methods, and results

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摘要 Background:Mostcurrentapproachesinforestscienceandpracticerequireinformationaboutstructureandgrowthofindividualtreesratherthan-orinadditionto-sumandmeanvaluesofgrowthandyieldatforeststandlevelasprovidedbyclassicexperimentaldesigns.Byinventingthewheeldesign,Nelderprovidedthepossibilitytoturntotheindividualtreeasbasicinformationunit.Suchtrialsprovidevaluableinsightsintothedependencyofgrowthonstanddensityatparticularsites.Methods:Here,wepresentanextensionoftheoriginaldesignandevaluationbyNelder.(i)WeestablishedNelderwheelsalonganenvironmentalgradientthroughEuropeinatlanticclimateinBelgiumandGermany,MediterraneanclimateinItaly,continentalclimateinHungaryaswellasonhighlandclimateinMexico.SuchdisjunctNelderwheelsalonganenvironmentalgradientcanberegardedandanalysedasatwo-factordesignwiththefactorsofsiteconditionandstanddensity.(ii)WepresentanadvancedstatisticalapproachtoevaluatedensitydependentgrowthdynamicsoftreesplantedinformoftheNelderdesign,whichconsidersspatio-temporalautocorrelation.(iii)Weprovetheusefulnessofthemethodsinimprovingecologicaltheoryconcerningdensityrelatedproductivity,trade-offsbetweenfacilitationandcompetition,andallometricrelationsbetweensizevariables.Results:FirstevaluationsbasedonremeasuredNelderwheelsinoak(QuercusroburL.)showasizegrowthdifferentiationduringthefirstobservationperiod.Inparticular,heightgrowthisacceleratedunderhighercompetitionindicatingfacilitationeffects.Wedetectfurthermoreahighvariabilityinallometricrelations.Conclusions:Theproposeddesign,methods,andresultsarediscussedregardingtheirimpactonforestpractice,modelbuilding,andecologicaltheory.WeconcludethattheextendedNelderapproachishighlyefficientinprovidingcurrentlylackingindividualtreelevelinformation.
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出版日期 2015年03月13日(中国期刊网平台首次上网日期,不代表论文的发表时间)